After watching Eljay's Recap Reviews of the Stars, I went on a nostalgia trip and re-played the Agori Defender game, which I could never beat when I was younger....and I beat it and was greeted with an interesting end cutscene featuring...a flying Tarduk?!
So the "spikes" on his shoulders (which I thought were defensive adaptations) were wings all along?
No wonder his legs are so thin! He never walks!