Skrall have consistently been referred to as members of the Rock Tribe, which they share with Agori of rock like Atakus. Does this mean that they are a subset of the Glatorian species, or are they a species all their own?
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Tribe membership doesn’t delineate species; Gresh is of the Jungle tribe as a Glatorian, as is the Agori Tarduk as a civilian. Not only are tribes trans-species coalitions, but Skrall are on numerous occasions specified to be a different species to the Glatorian. You can see this on BS01’s page on sapient species
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No, the Skrall are not of the Glatorian species. However, there are also no “subsets” of the Glatorian species. There are no genetic differences between a Glatorian-species of the Jungle Tribe vs. Water Tribe.
There might be regional differences, but that was never really covered in-story.
Now I’m wondering if any members of the Skrall species ever joined another Tribe.
The Rock Tribe probably wouldn’t let in any non-Skrall (at least not pre-Battle of Bara Magna), but I could see, for example, the Jungle Tribe welcoming a Skrall (with some suspicion).
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They are related to the main Glatorian species, but that’s about all we know.
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They are their own species. The Skrall are closely related to the Glatorian.